The subject of the sentence is "any mention of throughput or timing guarantees." It's a case where, when people try to sound smarter OR want to prequalify something, they place the syntax in a non-standard order.
The sentence essentially reads:
The mention of throughput and timing guarantees, which are services not provided by todays Internet transport protocols, was conspicuously missing in our brief description of TCP and UDP
For the second part, we have "conspicuously," meaning "obviously" or "clearly;" "throughput," or "amount passing through a system or process" or "what passes through the process;" and "timing guarantees," which research shows is the "amount of time a website is active."
The phrase as a whole says that
"Information on the amount of data passing through the system and the amount of uptime of a website are obviously missing"
The dash in front of "guarantees" indicates that this was a pause, and that the following phrase was describing the prior two.
As such, the whole sentence reads:
"Information on the amount of data passing through the system and the amount of uptime of a website, which are services not provided by today's Internet transport protocols, was obviously missing in our short description of TCP and UDP"
Or, more succinctly:
"Information on data transportation and website uptime was obviously not in our description of TCP and UDP, and these services are not provided by Internet Transport protocols"