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Similar to ROS, reactive nitrogen species (RNS) are by-product of cellular metabolism, primarily generated from mitochondria, and plays a fundamental role in aging

When I am typing this sentence, it occurs to me whether this past participle structure "primarily generated from mitochondria" here will cause ambiguity? I don't know normally how native English speakers use this structure. Is it more typical to write like this:

Similar to ROS, reactive nitrogen species (RNS) are by-product of cellular metabolism which are primarily generated from mitochondria, and plays a fundamental role in aging

Thanks!

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    The second sentence does make it more clear. Meanwhile, I would have almost said, "a by-product" or "by-products." – Teacher KSHuang Mar 16 '17 at 11:34
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    The verb plays should be in the plural (play), like are. – Ronald Sole Mar 16 '17 at 11:37
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    I would make two sentences. "... is a by-product of metabolism, generated primarily from mitochondria. RNS plays a fundamental role..." – Tᴚoɯɐuo Mar 16 '17 at 11:38
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    @RonaldSole I agree. 'play' refers to RNS! – SovereignSun Mar 16 '17 at 14:00

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