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the tough tissues surrounding the actual brain -- as well as tiny capillaries and portions of adjacent cortical tissues have been preserved as mineralised 'ghosts'. What is the meaning of 'ghosts' here ?

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https://www.merriam-webster.com/dictionary/ghost

Look at definition 4.a: "a faint shadowy trace."

The way a person leaves behind a ghost, the dinosaur's brain tissue left behind a fossil.

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