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Does the following sentence, mean the "ancestral communities" of A and B are exactly the same (bidirectional) or does it mean that only all "ancestral communities" of A are also "ancestral communities" of B, but B may have more "ancestral communities" (unidirectional)? If the structure is bidirectional, what other structure can I use to convey the unidirectional meaning?

If user A shares the same ancestral communities with user B, ...

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I think the phrase is bidirectional in itself. If you would like to convey the meaning that B may have more "ancestral communities", you should add a second sentence to provide more information on that matter.

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