I really couldn't decide on the meaning of "comprehend" in this sentence. Does it mean "understand" or "include/comprise/encompass", or something else? Could anyone help please? Many thanks.
He then told her of Georgiana’s delight in her acquaintance, and of her disappointment at its sudden interruption, which naturally leading to the cause of that interruption, she soon learnt that his resolution of following her from Derbyshire in quest of her sister had been formed before he quitted the inn, and that his gravity and thoughtfulness there had arisen from no other struggles than what such a purpose must comprehend.
from Pride & Prejudice