This is a rather convoluted sentence, but I think it breaks down like this:

• The obvious paradox which makes [the thesis] ... actually serve idealism ...
• the thesis of man’s animal origin
• the thesis thus understood
• [the paradox] has already been pointed out by A. Leroi-Gourhan.

So the part you've picked out is qualifying the thesis - it is not just any thesis of man's animal origins, it is the particular thesis that is understood because of something in a preceding sentence.

A more direct phrasing might be something like this:

An obvious paradox in this thesis of man's animal origins makes it actually serve idealism, as already pointed out by A. Leroi-Gourhan.

This is a rather convoluted sentence (you can read the entire article here), but I think it breaks down like this:

• The obvious paradox which makes [the thesis] ... actually serve idealism ...
• the thesis of man’s animal origin
• the thesis thus understood
• [the paradox] has already been pointed out by A. Leroi-Gourhan.

So the part you've picked out is qualifying the thesis - it is not just any thesis of man's animal origins, it is the particular thesis that is understood because of something in a preceding sentence.

A more direct phrasing might be something like this:

An obvious paradox in this thesis of man's animal origins makes it actually serve idealism, as already pointed out by A. Leroi-Gourhan.