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A correct squat perfectly balances all the forces around the knees and the hips, using these muscles in exactly the way the skeletal biomechanics are designed for them to be used, over their full range of motion.

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    across or spanning - in this exercise, these muscles go to 100% of their capacity (fully extended/contracted) and back to their starting position. Accordingly, your legs go from standing to fully bended in the knees.
    – CowperKettle
    Commented Jan 6, 2019 at 13:08

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The preposition over is often used to mean throughout or acting across. Thus, "over their full range of motion" means that the muscles are using their full range, stretching from one end/extreme to the other.

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